Gel Electrophoresis
In a native PAGE, proteins are separated on the basis of

net charge and size
net positive charge
net negative charge
net positive charges size

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Gel Electrophoresis
Electrophoresis of histones and myoglobin under non-denaturing conditions (pH = 7.0) results in

both proteins migrate to the anode
both proteins migrate to the cathode
histones migrate to the cathode and myoglobin migrates to the anode
histones migrate to the anode and myoglobin migrates to the cathode

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